Startseite Proportional population models and stability analysis on time scales
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Proportional population models and stability analysis on time scales

  • Ayşe Çiğdem Yar und Emrah Yilmaz EMAIL logo
Veröffentlicht/Copyright: 16. Juli 2025
Journal of Applied Analysis
Aus der Zeitschrift Journal of Applied Analysis

Abstract

The study of changes in population size and reasons for these changes appear as population dynamics. These dynamics generally focus on change in population size over time. Mathematical modeling is used to make sense of these changes. In this study, three population models have been reconstructed and solved using proportional derivative on time scales. Equilibrium points and stability states of these solutions were examined.

MSC 2020: 34N05; 26A33; 37C20

1 Introduction

A time scale 𝕋 is an arbitrary, nonempty, closed subset of ℝ. This theory was first introduced by Hilger [15]. It has been studied in detail by Bohner and Peterson [10, 11]. Additionally, fractional calculus is defined as a generalization of non-integer order ordinary derivatives and integrals [21]. Time scale and fractional theories were first combined by Bastos [7]. Proportional derivative was first studied on 𝕋 by Anderson and Ulness [5]. Anderson and Ulness discussed important definitions and theorems of proportional theory on 𝕋. In addition, Anderson and Georgiev [4] studied proportional dynamic equations on 𝕋 and gave important definitions and theorems. Now, let us remember some basic concepts of time scales.

According to [10], the forward jump operator σ : T T is σ ( t ) = inf { s T : s > t } for t T ; the graininess function μ : T [ 0 , ) is μ ( t ) = σ ( t ) t . Let f : T R be a function and t T κ . Then one can define f Δ ( t ) to be a number with the property that, given any ε > 0 , there is neighborhood 𝑈 of 𝑡 such that

| [ f ( σ ( t ) ) f ( s ) ] f Δ ( t ) [ σ ( t ) s ) ] | ε | σ ( t ) s | for all s U ,

where f Δ ( t ) is the delta derivative of 𝑓 at 𝑡.

A function f : T R is called regulated if it has finite right-sided limits at all right-dense points of 𝕋 and finite left-sided limits at all left-dense points of 𝕋. Let 𝑓 be regulated. Then there exists a pre-differentiable function 𝐹 with of differentiation region 𝐷 when F Δ ( t ) = f ( t ) holds for all t D . The indefinite delta integral of 𝑓 is f ( t ) Δ t = F ( t ) + C , where 𝐶 is an arbitrary constant.

Let h > 0 . The cylinder transformation ξ h : C h Z h is defined by

ξ h ( z ) = 1 h Log ( 1 + z h ) ,

where Log denotes the principal logarithm function. For h = 0 , the transformation is defined as ξ 0 ( z ) = z for all z C .

If p R , where ℛ is set of all regressive and rd-continuous functions, then the exponential function is

e p ( t , s ) = exp ( s t ξ μ ( τ ) ( p ( τ ) ) Δ τ ) for s , t T ,

where ξ μ ( τ ) is the cylinder transformation on 𝕋.

Let f : Λ n R and let t Λ i κ i n . Suppose that the limit lim s i t i f ( t s i ) = f ( t ) exists. If t i < σ i ( t i ) , then the function 𝑓 is said to be delta differentiable with respect to t i at 𝑡, and its delta derivative is given by (see [9])

f t i Δ i ( t ) = f t i σ i ( t ) f ( t ) μ i ( t i ) .

Population models are built upon birth, death, and migration rates. These models illustrate how various processes shape the size, composition, and dynamics of a population, including growth, decline, concentration, and renewal [24]. Schoen discusses numerous population models and mathematically analyzes features such as convergence to equilibrium, dynamic death rates, population structure, growth, and distribution. One of the most significant contributions to population modeling was made by Malthus, who introduced exponential growth based on a constant rate [18]. Following this study, many populations have been modeled using differential equations. Another notable population model was proposed by Verhulst [28], who incorporated Malthus parameters into his model [22]. Population modeling has been applied to various domains, including studies on exponential growth, logistic growth, interspecific competition, and real population estimates [23, 25, 19]. There are some recent studies in the literature that examine the dynamics of biological and physical systems using fractional calculus. Ahmed and Almatraf studied the stability of a discrete-time predator-prey model [1]. Berkal and Almatraf discussed the stability properties of a two-dimensional activator-inhibitor system with fractional order derivatives [8]. Khan and Almatraf studied the local dynamics and bifurcation behavior of a two-dimensional discrete-time laser model [17]. Almatrafi and Berkal investigated bifurcation analysis for the fractional predator-prey model with prey population growth [2]. Almatrafia and Messaoud Berkal studied the stability and bifurcation analysis of the predator-prey model they considered using conformable derivatives [3].

In our study, the Malthusian model will be discussed first. Malthus showed that the population tends to increase geometrically after a certain period of time [6]. This equation is

(1.1) d N d t = r N ( t ) ,

where N = N ( t ) represents number of individuals in a population and 𝑟 represents intrinsic growth rate [22]. Here r = b d , where 𝑏 is birth rate, 𝑑 is death rate, and 𝑟 is carrying capacity indicating maximum population size that a particular environment will support. The variable 𝑡 can be considered as year or month, depending on population characteristics considered.

Another population model we will consider in this study is the restricted two living state uniform growth model. This model is defined by [24] as

(1.2) d 1 d t = m 12 ( t ) 1 ( t ) + m 21 ( t ) 2 ( t ) , d 2 d t = m 21 ( t ) 2 ( t ) + m 12 ( t ) 1 ( t ) .

Here j is defined as number of people in state 𝑗 at time 𝑡 and m i j is defined as instantaneous transfer rate (or force) from state 𝑖 to state 𝑗 at 𝑡. This allows for movement between two states, but in both cases, the lack of growth is manifested by the sum of the above two equations always being zero. The reason for this is that this system of equations model allows transfer between states but assumes that there is no natural increase.

The last model we will discuss in this study is the interspecific competition model discussed by Lotka and Volterra [22]. The system of equations

(1.3) d N 1 d t = r 1 N 1 ( t ) ( 1 N 1 ( t ) + γ N 2 ( t ) K 1 ) , d N 2 d t = r 2 N 2 ( t ) ( 1 N 2 ( t ) + β N 1 ( t ) K 2 )

describes a classical competition model, where N 1 and N 2 represent the population size of two competitive populations, 𝛾, 𝛽, K 1 , K 2 , r 1 , r 2 are positive, and for i = 1 , 2 , the terms N i K i represent intraspecific competition. In this model, both types influence each other. Therefore, it is a symmetric system of equations. The carrying capacity 𝐾 becomes a shared resource, and γ , β are the pair of competition coefficients that define the degree of competition of each species over the other.

In this study, the three classical models given above will be defined and solved using proportional analysis on the time scales and stability analysis will be performed. The stability of solutions to a differential equation allows us to make predictions about how a process will unfold over time. From the perspective of differential equations, equilibrium points (or fixed points) represent equilibrium solutions. An equilibrium is defined as stable if all small disturbances fade away over time. Thus, stable equilibria are represented geometrically by stable fixed points. In contrast, in unstable equilibrium situations, disturbances grow over time [26]. One of the studies carried out in the process of examining population models is related to whether the processes have any steady state. In other words, whether the population increases, decreases, or remains constant over time is examined. Here the balance points are considered and the next population situation is examined assuming that the population amount changes after a fixed value [12]. Various studies have been conducted on this subject. Murray examined the stability of solutions in many population models [20]. Boyce and Diprima wrote a book explaining population dynamics and the stability of their solutions [13]. Stucchi et al. examined equilibrium solutions and stability of the solutions in a population dynamics model [27]. Salisbury carried out studies on equilibrium solutions and stability analysis by explaining many of the models [22].

This work is aimed at accurately modeling real world phenomena that exhibit the characteristics of both discrete and continuous states, such as population dynamics. Classical differential models often fail to account for discrete states in such systems. Proportional derivatives provide a more flexible modeling tool by generalizing the mathematical concepts used in mathematical modeling. Furthermore, combining proportional calculus with time scale theory provides a unified framework for addressing both discrete and continuous states within a single model. This approach not only increases the realism of the mathematical representation, but also demonstrates that approaches such as stability analysis used in classical analysis can also be expressed with the help of proportional derivatives. Therefore, the present work aims to contribute to the literature by establishing a proportional stability framework on time scales and demonstrating its effectiveness in analyzing nonlinear dynamical systems.

2 Preliminaries

In this section, basic theorems regarding stability analysis of equations and systems of equations in classical analysis are included. In addition, some important definitions and theorems regarding proportional dynamic equation theory are included in order to reconstruct the models using proportional theory.

Theorem 1

Theorem 1 ([22, 13])

Consider a differential equation, typically a nonlinear function of 𝑁,

d N d t = f ( N ) .

In this case, equilibrium points, where f ( N ) = 0 , are defined as N = N , where d N d t | N = N = 0 .

Theorem 2

Theorem 2 ([20])

For systems of equations, it is possible to find equilibrium points. Let 𝐴 be a 2 × 2 constant matrix and let 𝑥 be a 2 × 1 vector. Consider d x d t = A x . For this system, points where A x = 0 are equilibrium points.

Theorem 3

Theorem 3 ([16])

Let x = 0 be an equilibrium point for d x d t = f ( x ) , where f : D R n and D R n is a domain including x = 0 . Let V : D R be a continuously differentiable function where

V ( 0 ) = 0 and V ( x ) > 0 in D { 0 } , d V d t 0 in D .

Then x = 0 is stable (according to the Lyapunov criterion). Furthermore, if d V d t < 0 in D { 0 } , then x = 0 is asymptotically stable. Here V ( x ) is called a Lyapunov function.

Theorem 4

Theorem 4 ([14])

Consider the behavior of solutions for the autonomous system

(2.1) d x d t = f 1 ( x , y ) , d y d t = f 2 ( x , y )

near an isolated critical point ( x 0 , y 0 ) , where f 1 ( x 0 , y 0 ) = f 2 ( x 0 , y 0 ) = 0 . A critical point is isolated if its neighborhood includes no other critical point. Here f 1 and f 2 are continuously differentiable on a neighborhood of ( x 0 , y 0 ) . Let x 0 = y 0 = 0 without loss of generality. Otherwise, we make substitutions u = x x 0 , v = y y 0 . Then

d x d t = d u d t and d y d t = d v d t ,

so (2.1) is equivalent to

(2.2) d u d t = f 1 ( u + x 0 , v + y 0 ) = h 1 ( u , v ) , d v d t = f 2 ( u + x 0 , v + y 0 ) = h 2 ( u , v ) ,

that has an isolated critical point ( 0 , 0 ) .

Theorem 5

Theorem 5 ([14])

Taylor’s formula for two variable functions implies that if f 1 ( x , y ) is continuously differentiable near a fixed point ( x 0 , y 0 ) , then

f 1 ( x 0 + u , y 0 + v ) = f 1 ( x 0 , y 0 ) + f 1 x ( x 0 , y 0 ) u + f 1 y ( x 0 , y 0 ) v + r ( u , v ) .

If one applies Taylor’s formula to both f 1 and f 2 in (2.2) and supposes ( x 0 , y 0 ) is an isolated critical point,

f 1 ( x 0 , y 0 ) = f 2 ( x 0 , y 0 ) = 0 ,

this yields

(2.3) d u d t = f 1 x ( x 0 , y 0 ) u + f 1 y ( x 0 , y 0 ) v + r ( u , v ) , d v d t = f 2 x ( x 0 , y 0 ) + f 2 y ( x 0 , y 0 ) v + s ( u , v ) ,

where r ( u , v ) and the analogous remainder term s ( u , v ) for f 2 satisfy

(2.4) lim ( u , v ) ( 0 , 0 ) r ( u , v ) u 2 + v 2 = lim ( u , v ) ( 0 , 0 ) s ( u , v ) u 2 + v 2 = 0 .

Then, when 𝑢 and 𝑣 are small, r ( u , v ) and s ( u , v ) are small even in comparison with 𝑢 and 𝑣.

If one drops the presumably small nonlinear terms r ( u , v ) and s ( u , v ) in (2.3), the following linear system is obtained:

(2.5) d u d t = f 1 x ( x 0 , y 0 ) u + f 1 y ( x 0 , y 0 ) v , d v d t = f 2 x ( x 0 , y 0 ) + f 2 y ( x 0 , y 0 ) v ,

whose constant coefficients of u and v are values f 1 x ( x 0 , y 0 ) and f 1 y ( x 0 , y 0 ) of f 1 and f 2 at ( x 0 , y 0 ) . Because (2.3) is equivalent to the original nonlinear system u = f 1 ( x 0 + u , y 0 + v ) , v = f 2 ( x 0 + u , y 0 + v ) in (2.2), the conditions in (2.4) suggest that the linearized system in (2.5) closely approximates the given nonlinear system when ( u , v ) is close to ( 0 , 0 ) .

Supposing ( 0 , 0 ) is also an isolated critical point of the linear system, and the remainder terms in (2.3) satisfy the condition in (2.4), the original system x = f 1 ( x , y ) , y = f 2 ( x , y ) is said to be almost linear at ( x 0 , y 0 ) . So its linearization at ( x 0 , y 0 ) is a linear system in (2.5). Briefly, this linearization is the linear system u = J u (where u = [ u v ] T ) whose coefficient matrix is the Jacobian matrix

J ( x 0 , y 0 ) = [ f 1 x ( x 0 , y 0 ) f 1 y ( x 0 , y 0 ) f 2 x ( x 0 , y 0 ) f 2 y ( x 0 , y 0 ) ]

of f 1 and f 2 , evaluated at ( x 0 , y 0 ) .

Theorem 6

Theorem 6 ([14])

Let λ 1 and λ 2 be eigenvalues of the coefficient matrix of

(2.6) d x d t = a x + b y , d y d t = c x + d y

for a d b c 0 , associated with the almost linear system

(2.7) d x d t = a x + b y + r ( x , y ) , d v d t = c x + d y + s ( x , y ) .

  1. If λ 1 = λ 2 are real eigenvalues, then the critical point ( 0 , 0 ) of (2.7) is either a node or a spiral point, and is asymptotically stable if λ 1 = λ 2 < 0 , unstable if λ 1 = λ 2 > 0 .

  2. If λ 1 and λ 2 are pure imaginary, then ( 0 , 0 ) is either a center or a spiral point, and may be either asymptotically stable, stable, or unstable.

  3. Otherwise, the critical point ( 0 , 0 ) of the almost linear system in (2.7) is of the same type and stability as the critical point ( 0 , 0 ) of the associated linear system in (2.6).

The classical models discussed in this study will be addressed using proportional analysis on a time scale. Therefore, the following information should be given.

Definition 1

Definition 1 ([4])

Let α [ 0 , 1 ] and let k 0 , k 1 : [ 0 , 1 ] × T [ 0 , ) be continuous functions

lim α 0 + k 1 ( α , t ) = 1 , lim α 1 k 1 ( α , t ) = 0 , lim α 0 + k 0 ( α , t ) = 0 , lim α 1 k 0 ( α , t ) = 1 ,
k 1 ( α , t ) 0 , α [ 0 , 1 ) , k 0 ( α , t ) 0 , α ( 0 , 1 ] ,
where t T . If 𝑓 is Δ-differentiable at t T κ , the proportional Δ-derivative of 𝑓 at 𝑡 is

D α f ( t ) = k 1 ( α , t ) f ( t ) + k 0 ( α , t ) f Δ ( t ) .

Definition 2

Definition 2 ([4])

The proportional Δ-integral of 𝑓 on [ a , b ] is

a t f ( s ) Δ α , t s = a t f ( s ) E 0 ( t , σ ( s ) ) k 0 ( α , s ) Δ s , t [ a , b ] ,

where a , b T , a < b , and k 0 ( α , t ) μ ( t ) k 1 ( α , t ) 0 , α ( 0 , 1 ] , t T .

Definition 3

A function f : T R is called a conformable regressive function if

k 0 ( α , t ) μ ( t ) k 1 ( α , t ) 0 and k 0 ( α , t ) + μ ( t ) ( f ( t ) k 1 ( α , t ) ) 0

for any α ( 0 , 1 ] and for any t T . The set of all conformable regressive functions on 𝕋 is denoted by R c .

Definition 4

Definition 4 ([4])

Let α ( 0 , 1 ] , p R c . For t , t 0 T , the proportional exponential function is

E p ( t , t 0 ) = e p k 1 k 0 ( t , t 0 ) .

Theorem 7

Theorem 7 ([4])

Let 𝑓 and 𝑔 be Δ-differentiable at t T κ . Then

  1. D α ( f + g ) ( t ) = D α f ( t ) + D α g ( t ) ,

  2. D α ( f g ) ( t ) = ( D α f ( t ) ) g ( t ) + f σ ( t ) ( D α g ( t ) ) k 1 ( α , t ) f σ ( t ) g ( t ) .

Other proportional differentiation rules and properties can be found in [4].

Theorem 8

Theorem 8 ([4])

Let u : T × T C be Δ differentiable with respect to the first variable or the second variable at ( t , s ) T × T . The proportional Δ-partial derivatives with respect to the first variable or the second variable at ( t , s ) are

D t α u ( t , s ) = k 1 ( α , t ) u ( t , s ) + k 0 ( α , t ) u t Δ ( t , s ) , D s α u ( t , s ) = k 1 ( α , t ) u ( t , s ) + k 0 ( α , t ) u s Δ ( t , s ) ,

respectively.

Theorem 9

Theorem 9 ([4])

Let h C r d 1 ( T ) and g R c be such that

z h σ E g σ ( . , 0 ) = g E g ( . , 0 ) , D α h z h h σ + ( z k 1 ) h σ k 1 h = 0 , h ( 0 ) = 1

for z H c ( h ) , where H c ( h ) includes all z R c for z k 1 R c and

k 0 + h σ z ( μ k 1 ) 0 .

Let f : T C be regulated. The proportional Laplace transform of 𝑓 on time scales is

L c ( f ) ( z ) = 0 f ( t ) h σ ( t ) E g σ ( t , 0 ) Δ α , t

for z D c ( f ) , where D c ( f ) includes z H c ( h ) when the improper integral exists.

Theorem 10

Theorem 10 ([4])

Let A R c and q : T R n be rd-continuous. Let also t 0 T , y 0 R n , and let k 0 I + μ A be invertible on 𝕋. Then

D α y ( t ) = A ( t ) y ( t ) + q ( t ) , y ( t 0 ) = x 0 ,

has a unique solution

y : T R n , y ( t ) = y 0 E A ( t , t 0 ) + t 0 t E B ( σ ( s ) , t ) q ( s ) Δ α , t s ,

where B = ( μ k 1 k 0 ) A ( k 0 I + μ A ) 1 .

Theorem 11

Theorem 11 ([4])

Let 𝐴 be an n × n constant matrix and A R c , where R c is the set of all proportional regressive functions on 𝕋, and t 0 T . Consider the system

(2.8) D α x ( t ) = A x ( t ) ,

where x = ( x 1 x 2 x n ) T , 𝑇 is transpose. Let λ , ξ be an eigenpair of 𝐴. Then

x ( t ) = E λ ( t , t 0 ) ξ , t T κ ,

is a solution of (2.8).

Theorem 12

Theorem 12 (Proportional Putzer algorithm [4])

Let A R c be an n × n constant matrix and t 0 T . If λ 1 , λ 2 , , λ n are eigenvalues of 𝐴, then

E A ( t , t 0 ) = k = 0 n 1 r k + 1 ( t ) P k ,

where

r ( t ) = ( r 1 ( t ) r n ( t ) )

is solution of

D α r = ( λ 1 0 0 0 1 λ 2 0 0 0 1 λ 3 0 0 0 0 λ n ) r , r ( t 0 ) = ( 1 0 0 0 ) , P 0 = I , P k + 1 = ( A λ k + 1 I ) P k , 0 k n 1 .

3 Main results

In this section, equation (1.1) and systems (1.2), (1.3) are reconstructed using proportional dynamics equation theory. Then, similar to [13, 16, 20], stability states of proportional solutions are examined.

Theorem 13

The solution to the Malthusian proportional model

(3.1) D α N ( t ) = r N ( t ) ,
N ( 0 ) = β
is

(3.2) N ( t ) = β E r ( t , 0 ) ,

where β , r R .

Proof

Let us apply the proportional Laplace transform to both sides of (3.1) to get

L c { D α N } ( z ) = L c { r N } ( z ) = r L c { N } ,

and so

L c { N } ( z ) = β E 0 ( inf , 0 ) z r .

If the inverse proportional Laplace transform of both sides is taken from here, we get the solution of the Malthusian problem as

N ( t ) = L 1 { β E 0 ( inf , 0 ) z r } = β E r ( t , 0 ) ,

where L c 1 is called inverse proportional Laplace transform, which satisfies the property L c 1 ( F ) ( z ) = f ( . , 0 ) with L c ( f ( . , 0 ) ) ( z ) = F ( z ) . ∎

Conclusion 1

The only equilibrium point of equation (3.1) is N = 0 when D α N = 0 . This equation can show both exponential growth and exponential decreasing behavior. According to the definition of the proportional exponential function E r , exponential growth occurs when r > 0 , leading to unbounded growth. In this case, solution (3.2) is unstable. Conversely, when r < 0 , N ( t ) decreases over time and converges to zero. In this case, solution (3.2) is stable.

Theorem 14

Consider the restricted two living state uniform growth model

(3.3) D α 1 = m 12 ( t ) 1 ( t ) + m 21 ( t ) 2 ( t ) , D α 2 = m 21 ( t ) 2 ( t ) + m 12 ( t ) 1 ( t ) .

The solution of this system is

( t ) = c 1 E 0 ( t , t 0 ) ( m 21 ( t ) m 12 ( t ) 1 ) + c 2 E ( m 12 + m 21 ) ( t , t 0 ) ( 1 1 ) ,

where

A = ( m 12 ( t ) m 21 ( t ) m 12 ( t ) m 21 ( t ) ) .

Proof

Since

A = ( m 12 ( t ) m 21 ( t ) m 12 ( t ) m 21 ( t ) ) ,

we get

det ( m 12 ( t ) λ m 21 ( t ) m 12 ( t ) m 21 ( t ) λ ) = 0 .

Here λ 1 = 0 and λ 2 = ( m 12 + m 21 ) . Then the eigenvector corresponding to each eigenvalue will be found. For λ 1 = 0 , we obtain

( m 12 ( t ) m 21 ( t ) m 12 ( t ) m 21 ( t ) ) ( 1 2 ) = ( 0 0 ) and m 12 ( t ) 1 + m 21 ( t ) 2 = 0 , m 12 ( t ) 1 m 21 ( t ) 2 = 0 .

Thus, the eigenvector corresponding to the eigenvalue λ 1 is

ξ 1 = ( m 21 ( t ) m 12 ( t ) 1 ) .

Similarly, let us find the corresponding eigenvector λ 2 = ( m 12 + m 21 ) . Since

det ( m 21 ( t ) λ m 21 ( t ) m 12 ( t ) m 12 ( t ) λ ) = 0 ,

we get

m 21 ( t ) 1 + m 21 ( t ) 2 = 0 , m 12 ( t ) 1 m 12 ( t ) 2 = 0 .

So the eigenvector corresponding to λ 2 is

ξ 2 = ( 1 1 ) .

According to Theorem 11, the solution of system (3.3) is

( t ) = c 1 E 0 ( t , t 0 ) ( m 21 ( t ) m 12 ( t ) 1 ) + c 2 E ( m 12 + m 21 ) ( t , t 0 ) ( 1 1 ) .

Conclusion 2

Let us consider system (3.3). Let us find the equilibrium points 1 and 2 so that D α l 1 = 0 and D α l 2 = 0 , i.e.,

m 12 ( t ) 1 ( t ) + m 21 ( t ) 2 ( t ) = 0 1 = m 21 m 12 2 , m 21 ( t ) 2 ( t ) + m 12 ( t ) 1 ( t ) = 0 2 = m 12 m 21 1 .

When these two situations are considered together, the following is obtained:

1 = m 12 m 21 m 21 m 12 1 .

From here,

1 ( 1 m 12 m 21 m 21 m 12 ) = 0 .

Thus, we choose 1 = 0 and l 2 = 0 . So the equilibrium point is ( 1 , 2 ) = ( 0 , 0 ) .

Conclusion 3

Consider system (3.3). By Theorem 3, we choose

V ( l ) = 1 2 + 2 2 2 .

Then

D α V ( ) = 1 2 ( D α ( 1 2 ) + D α ( 2 2 ) ) , D α V ( ) = 1 2 ( m 12 ( 1 2 1 1 σ + 1 2 + 1 2 σ ) + m 21 ( 1 2 + 1 σ 2 2 2 2 2 σ ) k 1 ( 1 1 σ + 2 2 σ ) ) .

Here, if m 12 , m 21 , k 1 > 0 , 1 2 + 1 1 σ > 1 2 + 1 2 σ , 2 2 + 2 2 σ > 1 2 + 1 σ 2 , and 1 1 σ + 2 2 σ > 0 , we obtain

D α V ( ) 0 .

Thus, the ( 1 , 2 ) = ( 0 , 0 ) point is stable in the Lyapunov sense.

Theorem 15

Consider the interspecific competition model

(3.4) D α N 1 ( t ) = r 1 N 1 ( t ) ( 1 N 1 ( t ) + γ N 2 ( t ) K 1 ) , D α N 2 ( t ) = r 2 N 2 ( t ) ( 1 N 2 ( t ) + β N 1 ( t ) K 2 ) ,

and N ( t 0 ) = N 0 . The linearized form of the system around the equilibrium point ( N 1 , N 2 ) is given by

(3.5) D α u = [ k 0 ( α , N 2 ) r 1 N 1 μ ( N 2 ) ( 1 N 1 + γ σ ( N 2 ) K 1 ) ] u [ k 0 ( α , N 2 ) r 1 N 1 μ ( N 2 ) ( 1 N 1 + γ σ ( N 2 ) K 1 ) ] v , D α v = [ k 0 ( α , N 1 ) r 2 N 2 μ ( N 1 ) ( 1 N 2 + β σ ( N 1 ) K 2 ) ] u + [ k 0 ( α , N 2 ) r 2 N 2 μ ( N 2 ) ( 1 σ ( N 2 ) + β N 1 K 2 ) ] v .

Then the solution of the linearized system is expressed as

N ( t ) = ( 1 0 ) E A ( t , t 0 ) ,

where

N = ( u v ) , N 1 = β K 1 + K 2 β γ + 1 , N 2 = K 1 γ K 2 β γ + 1 , A = ( k 0 ( α , N 1 ) r 1 N 1 μ ( N 1 ) [ ( 1 σ ( N 1 ) + γ N 2 K 1 ) ] k 0 ( α , N 2 ) r 1 N 1 μ ( N 2 ) [ ( 1 N 1 + γ σ ( N 2 ) K 1 ) ] k 0 ( α , N 1 ) r 2 N 2 μ ( N 1 ) [ ( 1 N 2 + β σ ( N 1 ) K 2 ) ] k 0 ( α , N 2 ) r 2 N 2 μ ( N 2 ) [ ( 1 σ ( N 2 ) + β N 1 K 2 ) ] ) .

Proof

To linearize problem (3.4), the proportional Jacobian matrix

J ( N 1 , N 2 ) = ( D N 1 α ( r 1 N 1 ( 1 N 1 + γ N 2 K 1 ) ) D N 2 α ( r 1 N 1 ( 1 N 1 + γ N 2 K 1 ) ) D N 1 α ( r 2 N 2 ( 1 N 2 + β N 1 K 2 ) ) D N 2 α ( r 2 N 2 ( 1 N 2 + β N 1 K 2 ) ) )

is obtained, where

D N 1 α [ r 1 N 1 ( 1 N 1 + γ N 2 K 1 ) ] = k 1 ( α , N 1 ) [ r 1 N 1 ( 1 N 1 + γ N 2 K 1 ) ] + k 0 ( α , N 1 ) 1 μ ( N 1 ) [ r 1 N 1 ( 1 σ ( N 1 ) + γ N 2 K 1 ) r 1 N 1 ( 1 N 1 + γ N 2 K 1 ) ] ,
D N 2 α [ r 1 N 1 ( 1 N 1 + γ N 2 K 1 ) ] = k 1 ( α , N 2 ) [ r 1 N 1 ( 1 N 1 + γ N 2 K 1 ) ] + k 0 ( α , N 2 ) 1 μ ( N 2 ) [ r 1 N 1 ( 1 N 1 + γ σ ( N 2 ) K 1 ) r 1 N 1 ( 1 N 1 + γ N 2 K 1 ) ] ,
D N 1 α [ r 2 N 2 ( 1 N 2 + β N 1 K 2 ) ] = k 1 ( α , N 1 ) [ r 2 N 2 ( 1 N 2 + β N 1 K 2 ) ] + k 0 ( α , N 1 ) 1 μ ( N 1 ) [ r 2 N 2 ( 1 N 2 + β σ ( N 1 ) K 2 ) r 2 N 2 ( 1 N 2 + β N 1 K 2 ) ] ,
D N 2 α [ r 2 N 2 ( 1 N 2 + β N 1 K 2 ) ] = k 1 ( α , N 2 ) [ r 2 N 2 ( 1 N 2 + β N 1 K 2 ) ] + k 0 ( α , N 2 ) 1 μ ( N 2 ) [ r 2 N 2 ( 1 σ ( N 2 ) + β N 1 K 2 ) r 2 N 2 ( 1 N 2 + β N 1 K 2 ) ] .
From here,

J ( N 1 , N 2 ) = ( k 0 ( α , N 1 ) r 1 N 1 μ ( N 1 ) [ ( 1 σ ( N 1 ) + γ N 2 K 1 ) ] k 0 ( α , N 2 ) r 1 N 1 μ ( N 2 ) [ ( 1 N 1 + γ σ ( N 2 ) K 1 ) ] k 0 ( α , N 1 ) r 2 N 2 μ ( N 1 ) [ ( 1 N 2 + β σ ( N 1 ) K 2 ) ] k 0 ( α , N 2 ) r 2 N 2 μ ( N 2 ) [ ( 1 σ ( N 2 ) + β N 1 K 2 ) ] )

is obtained, where N 1 = β K 1 + K 2 β γ + 1 and N 2 = K 1 γ K 2 β γ + 1 are the equilibrium points of the problem. As a result, the linearized version of problem (3.4) is

D α u = [ k 0 ( α , N 2 ) r 1 N 1 μ ( N 2 ) ( 1 N 1 + γ σ ( N 2 ) K 1 ) ] u + [ k 0 ( α , N 2 ) r 1 N 1 μ ( N 2 ) ( 1 N 1 + γ σ ( N 2 ) K 1 ) ] v , D α v = [ k 0 ( α , N 1 ) r 2 N 2 μ ( N 1 ) ( 1 N 2 + β σ ( N 1 ) K 2 ) ] u + [ k 0 ( α , N 2 ) r 2 N 2 μ ( N 2 ) ( 1 σ ( N 2 ) + β N 1 K 2 ) ] v .

Then, by applying Theorem 10, we obtain

N ( t ) = ( 1 0 ) E A ( t , t 0 ) ,

where

N ( t ) = ( u v ) , N 1 = β K 1 + K 2 β γ + 1 , N 2 = K 1 γ K 2 β γ + 1 , A = ( k 0 ( α , N 1 ) r 1 N 1 μ ( N 1 ) [ ( 1 σ ( N 1 ) + γ N 2 K 1 ) ] k 0 ( α , N 2 ) r 1 N 1 μ ( N 2 ) [ ( 1 N 1 + γ σ ( N 2 ) K 1 ) ] k 0 ( α , N 1 ) r 2 N 2 μ ( N 1 ) [ ( 1 N 2 + β σ ( N 1 ) K 2 ) ] k 0 ( α , N 2 ) r 2 N 2 μ ( N 2 ) [ ( 1 σ ( N 2 ) + β N 1 K 2 ) ] ) .

Now let us calculate the E A ( t , t 0 ) function using the proportional Putzer algorithm for the special values we will consider for the linearized system.

Conclusion 4

Let k 0 = 2 , k 1 = 1 , γ = 4 , β = 6 , r 1 = 10 , r 2 = 10 , K 1 = 14 , K 2 = 2 , and T = Z . With the chosen parameter values, the equilibrium point is calculated as N 1 = 4 23 and N 2 = 7 23 . Accordingly, the point ( 4 23 , 7 23 ) is taken as the non-trivial equilibrium of the system. Since the system of equations (3.5) is a linearized system of equations, we need to look at its eigenvalues, det ( A I λ ) = 0 and λ 1 12.4348 , λ 2 19.5804 . From Theorem 12, the following system is obtained:

D α p = ( λ 1 0 1 λ 2 ) p , p ( t 0 ) = ( 1 0 ) .

So D α p 1 = λ 1 p 1 and D α p 2 = p 1 + λ 2 p 2 . By performing the necessary operations,

p 1 ( t ) = E λ 1 ( t , t 0 ) and p 2 ( t ) = t 0 t E λ 1 ( s , t 0 ) E g ( σ ( s ) , t ) Δ α , t s ,

where

g = ( 19.5804 k 1 ( α , t ) ) ( μ k 1 ( α , t ) ) k 0 ( α , t ) k 0 ( α , t ) + μ ( 19.5804 k 1 ( α , t ) ) ,

are obtained. On the other hand, since P k + 1 = ( A λ k + 1 I ) P k , 0 k n 1 , we obtain

P 0 = ( 1 0 0 1 ) , P 1 ( 17.4632 12.5520 20.2457 14.5520 ) .

From Theorem 12,

E A ( t , t 0 ) = p 1 ( 1 0 0 1 ) + p 2 ( 17.4632 12.5520 20.2457 14.5520 ) ,

and

( u v ) = ( c 1 c 2 ) ( p 1 ( 1 0 0 1 ) + p 2 ( 17.4632 12.5520 20.2457 14.5520 ) ) .

Let u ( t ) and v ( t ) denote the population growth functions of the first and second species, respectively. For the initial time t 0 = 0 , the population sizes at t = 3 months are calculated as u ( 3 ) 3474.93 , v ( 3 ) 1423.00 . These results suggest that the second species showed significant growth in the third month, while the first species exhibited a biologically unrealistic behavior by reaching a negative population size.

The equilibrium points of the solution obtained above are calculated as follows.

Conclusion 5

Let k 0 = 2 , k 1 = 1 , γ = 4 , β = 6 , r 1 = 10 , r 2 = 10 , K 1 = 14 , K 2 = 2 and T = Z . In this case, the linearized system (3.5) is analyzed around the equilibrium point ( N 1 , N 2 ) = ( 3.5652 , 0.2609 ) as follows. Since det ( J ( 0 , 0 ) I λ ) = 0 , we get

det ( J ( 3.5652 , 0.2609 ) I λ ) = 0 λ 1 42.2718 , λ 2 82.4549 .

Since both eigenvalues are real and positive, the equilibrium point is classified as an unstable improper node. This indicates that the system exhibits an unstable behavior around the equilibrium point and the solutions diverge from it over time.

4 Conclusions

In this study, classical population models were reconstructed and solved using the proportional derivative on time scales. The equilibrium points of these three models were calculated and an interpretation was made about the stability of the points. Stability analysis is used to qualitatively examine the behavior of the solutions we have considered. The method is based on determining the equilibrium points of the system, performing the proportional linearization via the proportional Jacobian matrix, and analyzing the resulting eigenvalue problem. This framework allows the study of the system behavior around equilibrium without requiring explicit solutions. Additionally, Lyapunov-based stability criteria are employed using proportional derivatives. The usage of proportional derivatives provides a flexible and unifying framework that generalizes classical and fractional models, capturing the system dynamics more accurately. By combining time scale theory with proportional calculus, the proposed method provides a robust and analytically tractable tool for studying stability in a wide range of applications.

References

[1] R. Ahmed and M. B. Almatrafi, Complex dynamics of a predator-prey system with Gompertz growth and herd behavior, Int. J. Anal. Appl. 21 (2023), 10.28924/2291-8639-21-2023-100. 10.28924/2291-8639-21-2023-100Suche in Google Scholar

[2] M. B. Almatrafi and M. Berkal, Bifurcation analysis and chaos control for fractional predator-prey model with Gompertz growth of prey population, Modern Phys. Lett. B 39 (2025), no. 23, Article ID 2550103. 10.1142/S0217984925501039Suche in Google Scholar

[3] M. B. Almatrafi and M. Berkal, Stability and bifurcation analysis of predator-prey model with Allee effect using conformable derivatives, J. Math. Comput. Sci. 36 (2025), no. 3, 299–316. 10.22436/jmcs.036.03.05Suche in Google Scholar

[4] D. R. Anderson and S. G. Georgiev, Conformable Dynamic Equations on Time Scales, CRC Press, Boca Raton, 2020. 10.1201/9781003057406Suche in Google Scholar

[5] D. R. Anderson and D. J. Ulness, Newly defined conformable derivatives, Adv. Dyn. Syst. Appl. 10 (2015), no. 2, 109–137. Suche in Google Scholar

[6] N. Bacaër, A Short History of Mathematical Population Dynamics, Springer, London, 2011. 10.1007/978-0-85729-115-8Suche in Google Scholar

[7] N. R. O. Bastos, Fractional calculus on time scales, Ph.D. Thesis, Instituto Politecnico de Viseu, 2012, Suche in Google Scholar

[8] M. Berkal and M. B. Almatrafi, Bifurcation and stability of two-dimensional activator-inhibitor model with fractional-order derivative, Fractal Fract. 7 (2023), no. 5, Paper No. 344. 10.3390/fractalfract7050344Suche in Google Scholar

[9] M. Bohner and S. G. Georgiev, Multivariable Dynamic Calculus on Time Scales, Springer, Cham, 2016. 10.1007/978-3-319-47620-9Suche in Google Scholar

[10] M. Bohner and A. Peterson, Dynamic Equations on Time Scales, Birkhäuser, Boston, 2001. 10.1007/978-1-4612-0201-1Suche in Google Scholar

[11] M. Bohner and A. Peterson, Advances in Dynamic Equations on Time Scales, Birkhäuser, Boston, 2003. 10.1007/978-0-8176-8230-9Suche in Google Scholar

[12] M. B. Bonsall, Population models, Ecology, Encyclopedia Life Support Syst. (EOLSS) 2 (2009), Paper No. 235. Suche in Google Scholar

[13] W. E. Boyce and R. C. DiPrima, Elementary Differential Equations and Boundary Value Problems, John Wiley & Sons, New York, 2009. Suche in Google Scholar

[14] C. H. Edwards and D. E. Penney, Differential Equations and Boundary Value Problems: Computing and Modeling, Pearson, London, 2000. Suche in Google Scholar

[15] S. Hilger, Ein Masskettenkalkül mit Anwendung auf Zentrumsmannigfaltigkeiten, Ph.D. Thesis, Universität Würzburg, 1988. Suche in Google Scholar

[16] H. K. Khalil, Control of Nonlinear Systems, Prentice Hall, New York, 2002. Suche in Google Scholar

[17] A. Q. Khan and M. B. Almatrafi, Two-dimensional discrete-time laser model with chaos and bifurcations, AIMS Math. 8 (2023), no. 3, 6804–6828. 10.3934/math.2023346Suche in Google Scholar

[18] T. Malthus, An Essay on the Principle of Population, J. Johnson, London, 1798. Suche in Google Scholar

[19] D. R. Mould and R. N. Upton, Basic concepts in population modeling, simulation, and model–based drug development, CPT Pharmacometrics Syst. Pharmacol. 1 (2012), no. 9, 1–14. 10.1038/psp.2012.4Suche in Google Scholar PubMed PubMed Central

[20] J. D. Murray, Mathematical Biology. I, Springer, New York, 2002. 10.1007/b98868Suche in Google Scholar

[21] K. B. Oldham and J. Spanier, The Fractional Calculus, Math. Sci. Eng. 111, Academic Press, New York, 1974. Suche in Google Scholar

[22] A. Salisbury, Mathematical Models in Population Dynamics, Thesis, Division of Natural Sciences New College of Florida, Sarasota, 2011. Suche in Google Scholar

[23] M. Schaub and F. Abadi, Integrated population models: A novel analysis framework for deeper insights into population dynamics, J. Ornithol. 152 (2011), 227–237. 10.1007/s10336-010-0632-7Suche in Google Scholar

[24] R. Schoen, Dynamic Population Models, Springer, New York, 2006. 10.1007/1-4020-5230-8Suche in Google Scholar

[25] R. M. Sibly and J. Hone, Population growth rate and its determinants: An overview, Philos. Trans. R. Soc. Lond. B Biol. Sci. 357 (2002), no. 1425, 1153–1170. 10.1098/rstb.2002.1117Suche in Google Scholar PubMed PubMed Central

[26] S. H. Strogatz, Nonlinear Dynamics and Chaos—With Applications to Physics, Biology, Chemistry, and Engineering, 3rd ed., CRC Press, Boca Raton, 2024. 10.1201/9780429398490Suche in Google Scholar

[27] L. Stucchi, J. M. Pastor, J. García-Algarra and J. Galeano, A general model of population dynamics accounting for multiple kinds of interaction, Complexity 2020 (2020), 1–14. 10.1155/2020/7961327Suche in Google Scholar

[28] P. F. Verhulst, Recherches mathématiques sur la loi d’accroissement de la population, Acad. Roy. de Bruxelles 18 (1845), 14–54. 10.3406/marb.1845.3438Suche in Google Scholar

Received: 2025-02-11
Revised: 2025-07-02
Accepted: 2025-07-03
Published Online: 2025-07-16

© 2025 the author(s), published by De Gruyter

This work is licensed under the Creative Commons Attribution 4.0 International License.

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